Mercury isotopes show vascular plants had colonized land extensively by the early Silurian.

The Hg isotope curve, however, does not show a decreasing trend before the Late Ordovician, indicating that, although their biomass may have been three times larger than today ( 21 , 22 ), the weathering flux resulting from nonvascular plant activity did not affect the Hg MIF signature observed in marine sediments.

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