Multi-trait genome-wide association analyses leveraging alcohol use disorder findings identify novel loci for smoking behaviors in the Million Veteran Program.

For example, one colocalized SNP for AUD and smoking initiation, rs6589386 on DRD2 , was marginally significant in the single-trait GWAS ( p -value = 7.17E-06) but attained GWS in MTAG ( p -value = 3.58E-10) (Fig. 3B ).

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