Origin of Low-Lying Red States in the Lhca4 Light-Harvesting Complex of Photosystem I.

Even though this shift is quite small (0.2 Å on average), its effect on the CT properties might be significant, given the sensitivity of CT couplings to the distance. 20 The mutual orientation between the chlorophylls is conserved among the WT replicas and with the mutant, leading to a well-defined and consistent orientation of the Chl pair.

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