The Eyes Absent family members EYA4 and EYA1 promote PLK1 activation and successful mitosis through tyrosine dephosphorylation.

Polo-like kinase 1 (PLK1) and Aurora kinase A (AURKA), two of the master regulatory kinases that control mitosis, were among the most highly significant EYA4 interacting proteins (Fig. 1A , Supplementary Data 1 ).

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