A distal enhancer guides the negative selection of toxic glycoalkaloids during tomato domestication.

To investigate whether the wrinkled surface phenotype of ws fruits is caused by the accumulation of phytotoxic SGAs compounds, we assessed SGAs content in fruit extracts by UPLC-MS revealing highly significant changes in γ-tomatine compared to other metabolites (Fig. 1e , Supplementary Fig. 3a–f ; Supplementary Data 5 ).

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