Upward and Poleward (but Not Phenological) Shifts in a Forest Tenebrionid Beetle in Response to Global Change in a Mediterranean Area.

velikensis changed their mean elevation from 393 m to 520 m (i.e., a 127 m uphill shift, which means an average shift of about 2 m per year), and the difference was marginally significant ( W = 9099, p = 0.060) ( Figure 3 A).

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