highly significantp < 2.2e-16
That is, traits within a group (e.g., both traits related to pigment level) are more highly significant (p < 2.2e-16 to 0.11, average p = 0.0041) than the significance is of correlations between groups (e.g., one trait is pigment level and another is patterning, p = 4e-13 to 0.99, average p = 0.148) ( S4 Table ).