Natural alleles of LEAFY and WAPO1 interact to regulate spikelet number per spike in wheat.

MADS-box transcription factors of the SQUAMOSA ( VRN1 and FUL2 ) and SVP clades ( SVP1 and VRT2 ) act as negative regulators of SNS, with loss-of-function mutations in any of these genes resulting in highly significant increases in SNS ( vrn1 58%, ful2 10%, vrt2 20%, and svp1 12%) (Li et al. 2019 , 2021 ).

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