Extreme Environmental Variability Induces Frontloading of Coral Biomineralisation Genes to Maintain Calcification Under pCO<sub>2</sub> Variability.

GO enrichment analysis was conducted for two WGCNA modules (MEbrown and MEred) with an expression pattern that showed highly significant positive correlation with net calcification and significant negative correlation with origin (i.e., the genes in these modules were higher expressed in the reef flat; Figure S5 ).

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