CHH hypermethylation contributes to the early ripening of grapes revealed by DNA methylome landscape of 'Kyoho' and its bud mutant.

Analysis of cluster 2 genes showed that methylation levels of ‘Fengzao’ sharply increased from F1 to F2, and then remained relatively invariable, whereas those of ‘Kyoho’ showed an increasing trend from K1 to K6 ( Fig. 4e ).

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On the other hand, 21 genes showed a decreasing trend during berry development, including DUF962 ( domain of unknown function 962 ), BURP ( BURP domain-containing protein ), EmrE ( multidrug resistance efflux transporter ), PME ( pectinesterase ), NXN ( nucleoredoxin ), TPT ( triose-phosphate transporter family ), UDPGT ( UDP-glucoronosyl and UDP-glucosyl transferase ), and RLK ( receptor-like serine/threonine-protein kinase ) ( Fig. 7a ).

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