Humerus shape evolved in cetaceans under relaxed selection and random drift.

Meanwhile, at the level of separate modules the marginal significance was found for proximal epiphysis ( p = 0.095) and diaphysis ( p = 0.054), also suggesting the weak negative selection, whereas in the distal epiphysis the expected and observed regression lines overlapped with a high support ( p = 0.769, Supplementary Table S10 ) supporting the neutral evolution of that structure.

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