an increasing trendP < 0.001
Long-term changes in iWUE and the effects of environmental variables on iWUE and ΔO 18 lw We observe an increasing trend in the iWUE of P. banksiana at all study sites: EST (slope = 0.45 μmol mol −1 year −1 , P < 0.001), ESB (slope = 0.682 μmol mol −1 year −1 , P < 0.001) and CLY (slope = 0.822 μmol mol −1 year −1 , P < 0.001) ( Figure 3c ).