Environmental change shapes understory plant diversity and dominance in boreal forests.

In terms of biomass proportion within the total understory community, increases in shrub biomass proportion decreased significantly with increasing cumulative nitrogen deposition ( P = 0.002), but showed a marginally significant increase in response to rising temperatures ( P = 0.069) (Fig. 3 d, Supplementary Fig. 3b , Supplementary Table 2 ).

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