highly significantP < 0.001
Compared with the null expectation of a constant frequency of innovation over time, the observed post-Ordovician decrease in the frequency of both unique and first occurrences of repeated phenotypes is highly significant (chi-square = 14.4 for unique traits, 43.6 for first occurrences of repeated traits, and 59.1 for the two groups combined, P < 0.001 in all cases).