marginally significantp = 0.056
In addition, we observed marginally significant negative indirect effects of LHC through PHYSCHEM (β = -0.125, p = 0.056) and VEG (β = -0.234, p = 0.061), suggesting a tendency for LHC to negatively impact the relative abundance of Pelophylax spp. by altering the vegetation and the physical and chemical characteristics of the waterbody (Online Resource Table 10 ).