highly significantp < 0.01
At the alpha diversity level, TN significantly positively affected TD (R = 0.28, p < 0.05 *), while TP exerted a highly significant positive effect on PD (R = 0.42, p < 0.01 **) and a highly significant negative effect on TD and FD (R = −0.42, p < 0.001 ***; R = −0.46, p < 0.001 ***).