highly significantP << 0.0001
Although there were no significant across population differences in mean observed heterozygosities (Ho) in both years (Friedman test: χ 2 0.05,5,17 = 5.662 , P = 0.340 for yr 1; yr 2 was similar), differences in allele composition and manner of pairing were evident from the highly significant genic and genotypic differentiation all P << 0.0001 .