A spreadable, non-integrative and high copy number shuttle vector for Sulfolobus solfataricus based on the genetic element pSSVx from Sulfolobus islandicus.

Interestingly, β-galactosidase activity in complemented mutant cells followed an increasing trend which correlated to the plasmid accumulation up to stationary phase and was not dependent on the number of the generations. lacS gene expression was inducible by thermal stress, namely it was confirmed to be transcriptionally regulated by the chaperonin gene promoter.

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