Transcription-related mutations and GC content drive variation in nucleotide substitution rates across the genomes of Arabidopsis thaliana and Arabidopsis lyrata.

These results were generated with gene density measured as the number of genes in a 0.50 Mb window centered around each intergenic locus; the result is similar, but only borderline significant, with 1.0 Mb windows ( r = -0.18; p = 0.09).

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