Genome-wide association studies identified three independent polymorphisms associated with α-tocopherol content in maize kernels.

Identification of Putative Causative and Independent Polymorphisms within the 2.4-Mb Region for Kernel αT Content Although the three ZmVTE4 SNPs showed highly significant associations with αT in CAM478 ( Figure 3B , Table S3 ), none of them were polymorphic in a By804/B73 recombinant inbred line (RIL) population, in which we identified a major QTL for αT near ZmVTE4 ( Figure S3 ) [21] .

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