Applying the 4 Population Test , we observed highly significant violations of the expected phylogenetic tree topology, confirming that the Roma are admixed; that is, they have ancestry from both South Asians and Europeans ( Table S2 ).
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Applying the 4 Population Test , we observed highly significant violations of the expected phylogenetic tree topology, confirming that the Roma are admixed; that is, they have ancestry from both South Asians and Europeans ( Table S2 ).