Comparing amphibians with reptiles, the largest upslope shifts occur with the frogs, especially the two microhylid genera Plethodontohyla and Platypelis , where all six species show a positive trend ( Fig. 4 , Table 1 ).
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Extinction vulnerability of tropical montane endemism from warming and upslope displacement: a preliminary appraisal for the highest massif in Madagascar
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These results show an overall trend for asymmetrical rates of invasion and contraction for both upslope and downslope movements, with 20 species exhibiting upslope shifts at the upper margin (range expansion) compared with 11 species exhibiting upslope shifts at the lower margin (range contraction).