marginal significanceP <0.10
Applying an FDR correction [39] using the program QVALUE (available from http://genomics.princeton.edu/storeylab/qvalue/ ) in the R statistics package ( http://www.r-project.org/ ) reduced this to ten loci at FDR <0.05, including four domestication-related and six improvement-related genes, with a three additional loci exhibiting marginal significance for selection during sunflower improvement after FDR correction (0.05< P <0.10).