Determinants and patterns of reproductive success in the greater horseshoe bat during a population recovery.

Among individual males, a steady increase in mean cumulative reproductive success was detected over the study period, which was also corroborated by the highly significant paternity skew estimated from the pooled paternity assignments based on 19 cohorts.

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A plot of cohort size against skew revealed a positive trend, illustrating that male reproductive skew among breeding males has tended to increase overall within the population between 1993 to 2011 ( Figure 2 ).

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