Non-denitrifying populations with a broad range of metabolisms and habitats may be significant contributors to the mitigation of N 2 O emissions. 5) Aerobic ammonia-oxidizing archaea ( Fig. 1 [5], Table 1 ) Ammonia oxidation, the first and rate-limiting step of nitrification, was considered to be performed mostly by certain groups of chemolithoautotrophic Proteobacteria (genera Nitrosospira , Nitrosomonas , and Nitrosococcus ) for more than one hundred years ( 44 ).
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