If a trait association was barely significant we conducted 1000 simulations in Mesquite with 10 iterations for the likelihood search to obtain a p value; for trait associations that were significant, we tested hypotheses about conditional evolution and the temporal order of trait acquisition by comparing the likelihood estimates of two correlated-evolution models: one in which all transition rates were allowed to vary and one that constrained the two transition rates to be equal (Figure 2 A).
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Correlates of monoicy and dioicy in hornworts, the apparent sister group to vascular plants.
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