Association mapping across numerous traits reveals patterns of functional variation in maize.

First, an analysis of expression levels using RNA-seq data from the Maize Gene Atlas [30] reveals a small (∼20%) but highly significant depletion of low-expressed genes (p = 1.30×10 −22 by Mann-Whitney test and ≈0 by Kolmogorov-Smirnov test) ( Fig. 6 ).

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