highly significantp<0.001
1 ) was highly significant (n = 204, r 2 = 0.951, p<0.001), the pattern and strength of relationship persisting when we controlled for phylogeny using two other family arrangements (99<n<134, 0.790< r 2 < 0.906, 493.46<F<969.27, p<0.001; Table S2).