Ecophenotypic Variation and Developmental Instability in the Late Cretaceous Echinoid Micraster brevis (Irregularia; Spatangoida).

Variation in non-morphometric characters Subanal fasciole, projection of the labrum, interradial structure of the paired petals, and granulation of the periplastronal area An obvious trend is seen in the development of the subanal fasciole, which is most pronounced in the Liencres population ( Fig 10A , S2 Table ), where a trace of a subanal fasciole is always present.

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