highly significantP < 0.001
However, the slope and strength of the relationship varied among species; feather δD increased with date in COYE ( b = 0.059 ± 0.025, r 2 = 0.025) and, more markedly, in WIWA ( b = 0.509 ± 0.106, r 2 = 0.148), but decreased with date in OCWA ( b = −0.154 ± 0.059, r 2 = 0.037) and YWAR ( b = −0.095 ± 0.037, r 2 = 0.033). For both plasma glycerol and β-hydroxybutyrate, there was a highly significant effect of year ( P < 0.001 in all cases; Table 1 ); however, the pattern of inter-annual variation was not consistent for these two metabolites.