marginally significantP = 0.096
Results The PERMANOVA including both species and FL factors identified significant differences in the elemental concentrations and their ratios and in the metabolomes of the needles of the three species (pseudo‐ F = 42.3; P < 0.0001), identified marginally significant differences between the FLs (pseudo‐ F = 1.9; P = 0.096) and identified significant differences in the species × FL interaction (pseudo‐ F = 2.7; P < 0.05) (Table 1 ).