highly significantP < 0.001
The PCA of shape variability from PC1 score showed that the forewing shape differences were highly significant among the 39 populations (F 2, 39 = 146.562, P < 0.001); however, the shape variability from PC2 score (F 2, 39 = 0.151, P > 0.05) and PC3 score (F 2, 39 = 0.010, P > 0.05) were not significant.