Beta-diversity analysis Average beta-diversity values were similar among years for the whole community (arithmetic mean ± SE: 0.419 ± 0.059 for 1997; 0.467 ± 0.070 for 2005; 0.444 ± 0.043 for 2012) and for the stygobiotic species (0.526 ± 0.065 for 1997; 0.632 ± 0.072 for 2005; 0.633 ± 0.049 for 2012; difference between 1997 and 2012 is in fact marginally significant), whereas increased significantly after the earthquake for the non-stygobites (0.340 ± 0.048 for 1997; 0.323 ± 0.044 for 2005; 0.535 ± 0.043 for 2012) (Table 3 ).
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Earthquake impacts on microcrustacean communities inhabiting groundwater-fed springs alter species-abundance distribution patterns.
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