Although it is conceivable that changes in scale accumulation rates could dilute TOC concentrations, the fact that no correlation exists with the total scale accumulation rate suggests that this is unlikely, and the ecological or habitat-based characteristics of hake may be significant at this site.
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Links between fish abundance and ocean biogeochemistry as recorded in marine sediments.
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The sentences
It seems quite feasible that upwelling-oxygen dynamics were responsible for the strong and highly significant negative correlations between TOC and Hake scales in the Soledad Basin.