highly significantP < 0.001
After incubation in the field, there was a highly significant effect of site ( F = 28, df = 1,32, P < 0.001; Appendix S1 : Table S2) and species of litter ( F = 26, df = 1,32, P < 0.001; Appendix S1 : Table S2) on the mass of carbohydrates remaining in litter, whereby this mass was lower in litter decomposing in forest plots and B. pubescens contained higher amounts of carbohydrates than E. nigrum , respectively (Fig. 3 a).