Broadcasting of amplitude- and frequency-modulated c-di-GMP signals facilitates cooperative surface commitment in bacterial lineages.

It is not known whether increased c-di-GMP levels constitute a necessary or sufficient condition for surface commitment: That increased c-di-GMP correlates with increased tendency for biofilm formation seems to be an overall trend that holds true, but there have also been many observed exceptions to this model, especially at the single-cell level.

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