A recent genome-wide association study (GWAS) has revealed that the number of spikelets on a panicle bears highly significant correlation with the number of secondary branches on it but only to a lesser extent with the number of primary branches ( Ta et al., 2018 ).
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Effect of Panicle Morphology on Grain Filling and Rice Yield: Genetic Control and Molecular Regulation.
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The reason for somewhat significant success, although a little, of both the programs was that these avoided the extremes of plant type traits ( Belford and Sedgley, 1991 ) and thus kept the targeted spikelet numbers per panicle to not more than 250 initially and later on not more than 150 spikelets per panicle.