We first extracted genotypes for the highly significant SNPs identified by GEMMA (13 for composite fitness, 4 for only growth: see section Genome-wide association tests ), and coded these as either reference, single, or double mutants using VCFtools ( Danecek et al., 2011 ).
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Hybridization alters the shape of the genotypic fitness landscape, increasing access to novel fitness peaks during adaptive radiation.
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