Overexpression of leucoanthocyanidin reductase or anthocyanidin reductase elevates tannins content and confers cassava resistance to two-spotted spider mite.

Furthermore, correlation analysis indicated that the transcription of tannin biosynthesis genes (i.e., F3H , DFR , ANR , ANS , and LAR ) and the abundance of most related metabolites (i.e., naringenin, gallocatechin, epigallocatechin, cyanidin, proanthocyanidin, catechin, epicatechin and procyandin B1) showed a positive correlation, particularly for LAR and ANR , which possessed the highest frequency of highly significant positive correlation ( Figure 2C ).

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