The relevance of these variables for explaining changes in mollusk species composition is demonstrated by the highly significant correlations between site scores on the first two PCoA axes (using the entire variance of the species data) (Supplementary Fig. 1 , Supplementary Table 3 ) and the CCA (using only the variance explained by the given factors) (Fig. 2 , Supplementary Table 2 ).
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The Holocene temperature conundrum answered by mollusk records from East Asia.
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