highly significantp < .001
A total of 153 highly significant ( p < .001) environmental variables were retained after forward selection, including 131 spatial variations, 10 energy variables (MAT, MDR, ISO, TS, MTWM, MTCM, MTWetQ, MTDQ, MTCQ, and WD), 8 significant water variables (MAP, PDM, PS, PWetQ, PDQ, PWarmQ, MI, and AET), 2 habitat heterogeneity‐related variables (ER and RMAP), and 2 human activity‐related variables (GDP and AOC) (Extended Data Table S2 ).