highly significantp < 0.01
Climatic variables Analysis of the environmental climatic variables using the Kruskal–Wallis test to compare the climatic niches of the two subspecies found two variables with highly significant differences ( p < 0.01; bio3 and bio4), two variables with significant differences ( p < 0.05; bio11 and bio18), and the other five variables were not statistically different ( p > 0.05; bio8, bio12, bio15, bio17, and bio18) ( Figures 4 , S5 ).