There were two additional SNPs on chromosomes 2 and 4 that approached significance (FDR = 0.11 and 0.14, respectively) and were therefore included for marker design (Fig. 2 ).
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Extreme-phenotype genome-wide association study (XP-GWAS) of powdery mildew race 2 W tolerance in the USDA Citrullus germplasm collection.
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