Genome-wide association study identifies novel variants in olfactory, vitamin A, vitamin B, and cadherin pathways associated with learning and memory.

In the GOBS cohort, proximate SNPs rs1032704 and rs4931672 at 12p11.21 were nominally significant for the Penn Conditional Exclusion Test (PCET) Face Memory Test and reached the suggestively significant threshold in the GRC GOM cohort for the SVLT LI measure for visual learning (Table 5).

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Set-based methods allow for the consideration of multiple SNPs within a gene that alone may not be highly significant but could have greater combined genetic effects 40 .

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