only marginally significantP = 0.066
The occupancy model, which separates detection probability from true occupancy, revealed a strong negative effect of road distance on occupancy (coefficient = -0.5897, SE = 0.0655, P< 0.0001), while the detection covariate (accessibility) was only marginally significant (P = 0.066) ( Supplementary Material S2 , Supplementary Table 2A ).