The excess of nominally significant association for flowering-related phenotypes has been eliminated, as would be expected if this excess were mostly due to confounding by population structure.
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Genome-wide association study of 107 phenotypes in Arabidopsis thaliana inbred lines.
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The sentences
There is a marked reduction in the number of associations of moderate significance (e.g., around 10 −4 ) across phenotypes, but the excess of highly significant associations clearly persists (or has even become greater).