Fungal communities of heavily invaded sites were shifted along NMDS axis 1 in relation to those of uninvaded and lightly invaded sites (Figure 4 C), but PerMANOVA found invasion level to be only marginally significant when soil texture and habitat were included in the model (Table 2 ).
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Soil Bacteria and Fungi Respond on Different Spatial Scales to Invasion by the Legume Lespedeza cuneata.
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