moderately significantp = 0.020
However, moderately significant linear relationships were detected between bleaching (loss of Chl a ) and chronic photodamage (reduced F v / F m ) in both H. depressa (R 2 = 0.532, p = 0.020) and M. vertebralis (R 2 = 0.300, p = 0.034) (data not shown).