Paleoceanographic insights on recent oxygen minimum zone expansion: lessons for modern oceanography.

PLoS One · 2015 · PMC4309540 · PMID 25629508

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However, this region exhibits a clear trend of higher oxygenation in the LGM, transitioning to severe hypoxia in intermediate waters at the start of the Holocene.

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Quoted from the open-access full text in Europe PMC under the licence the publisher applied. The sentence is reproduced exactly as published; the emphasis is ours.