The Surales, Self-Organized Earth-Mound Landscapes Made by Earthworms in a Seasonal Tropical Wetland.

This difference was not significant in Site 1 (two-way MANOVA, P > 0.05) but was highly significant in Sites 2 and 3 (two-way MANOVA, P < 0.001 in both sites).

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For example, in the mound phytolith profile in Site 1, panicoid grasses showed a decreasing trend toward the surface and in the mound phytolith profile in Site 2, they fluctuated with no clear increasing trend.

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In Sites 1 and 2, although grasses of this taxon were 2–15 times more abundant on mounds than in inter-mound basins, depending on site and season, no marked difference was apparent in the representation of phytolith morphotypes characteristic of panicoids in surface soils between these two habitats in either site ( Fig 4 ).

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